Microfibrils are synthesized by large membrane complexes that contain three isoforms of cellulose synthase, which are glucosyltransferases that are encoded by CESA genes.Ī membrane-bound endoglucanase, which is called KORRIGAN, is required for proper formation of the cellulose microfibril, perhaps serving a trimming function. The growing cell wall in plants is a thin, strong and pliant extracellular layer, composed of cellulose microfibrils embedded in a hydrated matrix that is made of complex polysaccharides and a small amount of structural proteins.Ī cellulose microfibril consists of ∼36 (1,4)-linked β- D-glucans that crystallize into a strong, thin ribbon ∼4 nm in diameter.
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